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List of publications  

Publications starting in 2000  

Publications starting in 2007  

Publications starting in 2013  

Publications starting in 2017  

Publications starting in 2020  

Non-technical publications  

 

WALTER D. KOENIG

Address          Hastings Reservation

                       38601 E. Carmel Valley Rd.                         

                        Carmel Valley, CA 93924

                        email: wdk4@cornell.edu

Born               1950, US citizen

                        Married to Janis L. Dickinson; 2 children

 

Education and job experience

 

2016-           Graduate School Professor, Field of Neurobiology and Behavior,

                        Cornell University

2008-            Research Zoologist Emeritus, Univ. California, Berkeley

2008-16           Senior Scientist, Cornell Lab of Ornithology and Department of

                        Neurobiology and Behavior, Cornell University

1982-2008           Research Zoologist, Museum of Vertebrate Zoology

                        University of California, Berkeley and Adjunct Professor

                        Department of Integrative Biology

1981-82           Assistant Professor and Curator, Moore Laboratory of Zoology,

                        Occidental College, Los Angeles, California

1978-81           Postdoctoral Research Zoologist, Museum of Vertebrate

                        Zoology, University of California, Berkeley

1972-78           Graduate Student, University of California, Berkeley.

                        Ph.D. in Zoology, December 1978

1968-72           Undergraduate, Stanford University, B.S. in Biology, June 1972

 

Academic and professional honors

 

2016    Lloye and Alden Miller Award, Cooper Ornithological Society

            (for citation, see The Condor 118: [2016])

2002    Honorary Member, Cooper Ornithological Society

            (for citation, see The Condor 105: 175 [2003])

1999    William Brewster Memorial Award, American Ornithologists’ Union

            (for citation, see The Auk 117: 265-266 [2000])

1998    Fellow, California Academy of Sciences

1998    Visiting Researcher, spatial ecology program, University of Helsinki

1988    Fellow, American Ornithologists' Union

1988    Fulbright Fellowship (Cambridge, UK)

1982    Elective Member, American Ornithologists' Union

 

Selected grants and fellowships

 

2015               NSF research grant (Evolution of cooperative behavior)

2009               NSF research grant (Reproductive fitness and sharing in cooperative                      societies)

2008, 2013     NSF research grant (LTREB: Phenology and masting in California oaks)

2006                NSF research grant (Ecological factors affecting group living)

2006                 Integrated Hardwoods Range Management Program, UC Berkeley  (Factors Affecting Valley and Blue Oak Regeneration in upper Carmel Valley, California)

1977-78           Annie M. Alexander Fellowship, Museum of Vertebrate Zoology

1975-78           NSF Predoctoral aid-to-dissertation Grant

1972-75           NSF Graduate Fellowship.

 

Major service positions

 

International Society for Behavioral Ecology, Secretary 1992-94; Treasurer 2000-2016

Organized biennial meeting, International Society for Behavioral Ecology, 1998

Council, American Ornithologists’ Union, 1999-2001

Editor, The Condor, 1995-2000

Associate Editor, Ecography, 2005-2009

Associate Editor, Ethology, 2012-

Associate Editor, Ecology/Ecological Monographs , 2005 - 2017

Board of Directors, Cooper Ornithological Society, 1992-1995

Editor, California Acorn Report, 1997 -

 

Fields of research

 

Reproductive strategies of forest trees; spatial synchrony; spatial patterns; conservation biology; behavioral ecology; ornithology; population ecology; vertebrate social behavior; dispersal, mating systems, and sexual selection.

 
 

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Publications starting in 2000  

Not-so-distant past publications (2007- )  

More-or-less current publications (2015- )  

Non-technical publications  

PUBLICATIONS OF WALTER D. KOENIG

 

Books

 

1. Koenig, W. D. and R. L. Mumme. 1987. Population Ecology of the Cooperatively Breeding Acorn Woodpecker. xiii+435 pp. Monographs in Population Biology 24, Princeton University Press, Princeton, NJ.

2. Stacey, P. B. and W. D. Koenig (editors). 1990. Cooperative Breeding in Birds: Long-term Studies of Ecology and Behavior. xviii+615 pp. Cambridge University Press, Cambridge.

3. Koenig, W. D. and J. L. Dickinson (editors). 2004. Ecology and Evolution of Cooperative Breeding in Birds. ix+293 pp. Cambridge University Press, Cambridge.

4. Koenig, W. D. and J. L. Dickinson (editors). 2016. Cooperative Breeding in Vertebrates: Studies of Ecology, Evolution, and Behavior. x+379 pp. Cambridge University Press, Cambridge.

 

Technical articles

 

1. Gutièrrez, R. J. and W. D. Koenig. 1978. Characteristics of storage trees used by acorn woodpeckers in two California woodlands. Journal of Forestry 76: 162-164.

 2. Koenig, W. D. and P. L. Williams. 1979. Notes on the status of acorn woodpeckers in central Mexico. Condor 81: 317-318.

 3. Koenig, W. D. and F. A. Pitelka. 1979. Relatedness and inbreeding avoidance: counterploys in the communally nesting acorn woodpecker. Science 206: 1103-1105.

 4. Koenig, W. D. 1980. The determination of runt eggs in birds. Wilson Bulletin 92: 103-107.

 5. Koenig, W. D. 1980. The incidence of runt eggs in woodpeckers.Wilson Bulletin 92: 169-176.

 6. Koenig, W. D. 1980. Variation and age determination in a population of acorn woodpeckers. Journal of Field Ornithology 51: 10-16.

 7. Davis, J., W. D. Koenig and P. L. Williams. 1980. Birds of Hastings Reservation, Monterey County California. Western Birds 11:113-128.

 8. Koenig, W. D. 1980. Acorn storage by acorn woodpeckers in an oak woodland: an energetics analysis. Pp. 265-269 in Symposium on the Ecology, Management, and Utilization of California Oaks (T. R. Plumb, technical coordinator) Pacific SW Forest & Range Exp. Station Gen. Tech. Rep. PSW44.

 9. Williams, P. L. and W. D. Koenig. 1980. Water dependence of birds in a temperate oak woodland. Auk 97: 339-350.

10. Koenig, W. D. 1981. Space competition in the acorn woodpecker: power struggles in a cooperative breeder. Animal Behaviour 29: 396-409.

11. Koenig, W. D. 1981. Reproductive success, group size, and the evolution of cooperative breeding in the acorn woodpecker. American Naturalist 117: 421-443.

12. Koenig, W. D., and F. A. Pitelka. 1981. Ecological factors and kin selection in the evolution of cooperative breeding in birds. Pp. 261-280 in Natural Selection and Social Behavior: Recent Research and New Theory (R. D. Alexander and D. W. Tinkle, editors) Chiron Press, New York.

13. Koenig, W. D. 1981. Coalitions of male lions: making the best of a bad job? Nature 293: 413. pdf

14. Koenig, W. D. 1982. Ecological and social factors affecting hatchability of eggs. Auk 99:526-536. pdf

Data from this paper: List of references (pdf) || Data summary (pdf)

15. Joste, N. E., W. D. Koenig, R. L. Mumme and F. A. Pitelka. 1982. Intragroup dynamics of a cooperative breeder: an analysis of reproductive roles in the acorn woodpecker. Behavioral Ecology & Sociobiology 11: 195-201.

16. Koenig, W. D., R. L. Mumme, and F. A. Pitelka. 1983. Female roles in cooperatively breeding acorn woodpeckers. Pp. 235-261 in Social Behavior of Female Vertebrates (S. K. Wasser, editor) Academic Press, New York.

17. Mumme, R. L., W. D. Koenig and F. A. Pitelka. 1983. Mate-guarding in the acorn woodpecker: within-group reproductive competition in a cooperative breeder. Animal Behaviour 31:1094-1106.

18. Mumme, R. L., W. D. Koenig and F. A. Pitelka. 1983. Are acorn woodpecker territories aggregated? Ecology 64: 1305-1307.

19. Mumme, R. L., W. D. Koenig and F. A. Pitelka. 1983. Reproductive competition in the communal acorn woodpecker: sisters destroy each other's eggs. Nature 306: 583-584.

20. Koenig, W. D. 1984. Clutch size of the gilded flicker. Condor 86: 89-90.

21. Koenig, W. D., R. L. Mumme and F. A. Pitelka. 1984. The breeding system of the acorn woodpecker in central coastal California. Zeitshrift für Tierpsychologie 65: 289-308.

22. Koenig, W. D. 1984. Geographic variation in clutch size in the northern flicker (Colaptes auratus ): support for Ashmole's hypothesis. Auk 101: 698-706.

23. Mumme, R. L., W. D. Koenig, R. M. Zink and J. A. Marten. 1985. An analysis of genetic variation and parentage in a California population of acorn woodpeckers. Auk 102: 305-312.

24. Koenig, W. D. and S. S. Albano.1985. Patterns of territoriality and mating success in the white-tailed skimmer Plathemis lydia (Odonata: Anisoptera). American Midland Naturalist 114: 1-12.

25. Koenig, W. D. 1985. Dunking of prey by Brewer's blackbirds: a novel source of water for nestlings. Condor 87: 444-445.

26. Hannon, S. J., R. L. Mumme, W. D. Koenig and F. A. Pitelka.1985. Replacement of breeders and within-group conflict in the cooperatively breeding acorn woodpecker. Behavioral Ecology & Sociobiology 17: 303-312.

27. Moore, W. S. and W. D. Koenig. 1986. Comparative reproductive success of yellow-shafted, red-shafted and hybrid flickers across a hybrid zone. Auk 103: 42-51.

28. Koenig, W. D. and S. S. Albano. 1986. On the measurement of sexual selection. American Naturalist 127: 403-409. pdf

29. Koenig, W. D. 1986. Geographic ecology of clutch size variation in North American woodpeckers. Condor 88: 499-504.

30. Hannon, S. J., R. L. Mumme, W. D. Koenig, S. Spon and F. A. Pitelka. 1987. Poor acorn crop, dominance, and decline in numbers of acorn woodpeckers. Journal of Animal Ecology 56: 197-207.

31. Koenig, W. D. and S. S. Albano. 1987. Lifetime reproductive success, selection, and the opportunity for selection in the white-tailed skimmer Plathemis lydia (Odonata: Libellulidae). Evolution 41: 22-36. pdf

32. Koenig, W. D. and M. D. Reynolds. 1987. Potential poisoning of yellow-billed magpies by compound 1080. Wildlife Society Bulletin 15: 274-276.

33. Koenig, W. D. and S. S. Albano. 1987. Breeding site fidelity in Plathemis lydia (Anisoptera: Libellulidae). Odonatologica 16: 249-259.

34. Carmen, W. J., W. D. Koenig and R. L. Mumme. 1987. Acorn production by five species of oaks over a seven year period at the Hastings Reservation, Carmel Valley, California. Pp. 429-434 in Symposium on Multiple-Use Management of California's Hardwood Resources (T. R. Plumb and N. H. Pillsbury, technical coordinators). Pacific SW Forest & Range Exp. Station Gen. Tech. Rep. PSW100.

35. Koenig, W. D. 1987. Morphological and dietary correlates of clutch size in North American woodpeckers. Auk 104: 757-765.

36. Koenig, W. D., S. J. Hannon, R. L. Mumme and F. A. Pitelka. 1988. Parent-offspring conflict in the acorn woodpecker. Proceedings of the International Ornithological Congress 19: 1221-1230.

37. Mumme, R. L., W. D. Koenig and F. A. Pitelka. 1988. Costs and benefits of joint nesting in the acorn woodpecker. American Naturalist 131: 654-677.

38. Koenig, W. D. 1988. Reciprocal altruism in birds: a critical review. Ethology and Sociobiology 9: 73-84. pdf

39. Koenig, W. D. 1988. On the determination of viable population size in birds and mammals. Wildlife Society Bulletin 16: 230-234.

40. Koenig, W. D. and M. K. Heck. 1988. Ability of two species of oak woodland birds to subsist on acorns. Condor 90: 705-708.

41. Koenig, W. D. 1988. Internal migration in the contemporary United States: comparison of measures and partitioning of stages. Human Biology 60: 927-944.

42. Koenig, W. D. 1989. Sex-biased dispersal in the contemporary United States. Ethology and Sociobiology 10: 263-278. pdf

43. Mumme, R. L., W. D. Koenig and F. L. W. Ratnieks. 1989. Helping behaviour, reproductive value, and the future component of indirect fitness. Animal Behaviour 38: 331-343.

44. Koenig, W. D. 1990. Territory size and duration in the white-tailed skimmer Plathemis lydia (Odonata: Libellulidae). Journal of Animal Ecology 59: 317-334.

45. Stacey, P. B. and Koenig, W. D. 1990. Introduction. Pp. ix-xviii in Cooperative Breeding in Birds: Long-term Studies of Ecology and Behavior (P. B. Stacey and W. D. Koenig, editors). Cambridge University Press, Cambridge.

46. Koenig, W. D. and P. B. Stacey. 1990. Acorn woodpeckers: Group living and food storage under contrasting ecological conditions. Pp. 415-453 in Cooperative Breeding in Birds: Long-term Studies of Ecology and Behavior (P. B. Stacey and W. D. Koenig, editors). Cambridge University Press, Cambridge.

47. Koenig, W. D. and R. L. Mumme. 1990. Levels of analysis and the functional significance of helping behavior. Pp. 268-303 in Interpretation and explanation in the study of behavior, vol. 2: Explanation, Evolution, and Adaptation (M. Bekoff and D. Jamieson, editors). Westview Press, Boulder, Colorado. Reprinted in: Readings in Animal Cognition (M. Bekoff and D. Jamieson, editors). MIT Press, Cambridge, Massachusetts.

48. Weathers, W. W., W. D. Koenig and M. T. Stanback. 1990. Breeding energetics and thermal ecology of the acorn woodpecker in central coastal California. Condor 92: 341-359.

49. Mumme, R. L., W. D. Koenig and F. A. Pitelka. 1990. Individual contributions to cooperative nest care in the acorn woodpecker. Condor 92: 360-368.

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50. Koenig, W. D. 1990. Dispersal, effective population size and the genetic structure of the contemporary United States. American Journal of Human Biology 2: 165-170. pdf

51. Koenig, W. D. 1990. Opportunity of parentage and nest destruction in polygynandrous acorn woodpeckers, Melanerpes formicivorus. Behavioral Ecology 1: 55-61.

52. Koenig, W. D., S. S. Albano and J. L. Dickinson. 1991. A comparison of methods to partition selection acting via components of fitness: do larger male bullfrogs have greater hatching success? Journal of Evolutionary Biology 4: 309-320. pdf

53. Koenig, W. D. 1991. The effects of tannins and lipids on the digestibility of acorns by acorn woodpeckers. Auk 108: 79-88.

54. Koenig, W. D., W. J. Carmen, M. T. Stanback and R. L. Mumme. 1991. Determinants of acorn productivity among five species of oaks in central coastal California. Pp.136-142 in Proceedings of the Symposium on Oak Woodlands and Hardwood Rangeland Management. (R. B. Standiford, Technical coordinator). Pacific SW Forest & Range Exp. Station Gen. Tech. Rep. PSW126.

55. Koenig, W. D., M. T. Stanback, P. N. Hooge and R. L. Mumme. 1991. Distress calls in the acorn woodpecker. Condor 93: 637-643.

56. Mumme, R. L. and W. D. Koenig. 1991. Explanations for avian helping behavior. Trends in Ecology & Evolution 6: 343-344.

57. Koenig, W. D. 1991. Levels of female choice in the white-tailed skimmer Plathemis lydia (Odonata: Libellulidae). Behaviour 119: 193-224.

58. Koenig, W. D., P. A. Gowaty and J. L. Dickinson. 1992. Boxes, barns, and bridges: confounding factors or exceptional opportunities in ecological studies? Oikos 63: 305-308.

59. Stanback, M. T. and W. D. Koenig. 1992. Cannibalism in birds. Pp. 277-298 in Cannibalism: Ecology and Evolution among Diverse Taxa (M. A. Elgar and B. J. Crespi, eds.). Oxford University Press, Oxford.

60. Koenig, W. D., F. A. Pitelka, W. J. Carmen, R. L. Mumme and M. T. Stanback. 1992. The evolution of delayed dispersal in cooperative breeders. Quarterly Review of Biology 67: 111-150. pdf

61. Eichholz, M. W. and W. D. Koenig. 1992. Gopher snake attraction to birds' nests. Southwestern Naturalist 37: 293-298. pdf

62. Birkhead, T. R., K. Clarkson, M. D. Reynolds and W. D. Koenig. 1992. Copulation and mate guarding in the colonial yellow-billed magpie Pica nuttalli and a comparison with the solitary black-billed magpie P. pica. Behaviour 121: 110-130.

63. Koenig, W. D., R. L. Mumme, W. J. Carmen and M. T. Stanback. 1994. Acorn production by oaks in central coastal California: variation within and among years. Ecology 75: 99-109. pdf

64. Stanback, M. T. and W. D. Koenig. 1994. Techniques for capturing birds inside natural cavities. Journal of Field Ornithology 65: 70-75.

65. Du Plessis, M., W. W. Weathers and W. D. Koenig. 1994. Energetic benefits of communal roosting by acorn woodpeckers in the non-breeding season. Condor 96: 631-637.

66. Leonard, M. L., K. L. Teather, A. G. Horn, W. D. Koenig and J. L. Dickinson. 1994. Provisioning in western bluebirds is not related to offspring sex. Behavioral Ecology 5: 455-459.

67. Knops, J. M. H. and W. D. Koenig. 1994. Water use strategies of five sympatric species of Quercus in central coastal California. Madroño 41: 290-301.

68. Koenig, W. D., J. M. H. Knops, W. J. Carmen, M. T. Stanback and R. L. Mumme. 1994. Estimating acorn crops using visual surveys. Canadian Journal of Forest Research 24: 2105-2112. pdf

69. Koenig, W. D. 1994. Two new bird-mammal associations from Kenya, with comments on host use by wattled starlings. Ostrich 65: 337-338.

70. Koenig, W. D., R. L. Mumme, M. T. Stanback and F. A. Pitelka. 1995. Patterns and consequences of egg destruction among joint-nesting acorn woodpeckers. Animal Behaviour 50: 607-621.

71. Dickinson, J. L., J. Haydock, W. D. Koenig, M. T. Stanback and F. A. Pitelka. 1995. Genetic monogamy in single-male groups of acorn woodpeckers. Molecular Ecology 4: 765-770.

72. Leonard, M. L., J. L. Dickinson, A. G. Horn and W. D. Koenig. 1995. An experimental test of offspring recognition in western bluebirds. Auk 112: 1062-1064.

73. Koenig, W. D., P. B. Stacey, M. T. Stanback and R. L. Mumme. 1995. Acorn woodpecker (Melanerpes formicivorus). In The Birds of North America, No. 194 (A. Poole and F. Gill, eds.) Academy of Natural Sciences, Philadelphia; American Ornithologists’ Union, Washington, D.C.

74. Dickinson, J. L., W. D. Koenig and F. A. Pitelka. 1996. Fitness consequences of helping behavior in the western bluebird. Behavioral Ecology 7: 168-177.

75. Koenig, W. D., J. M. H. Knops, W. J. Carmen, M. T. Stanback and R. L. Mumme. 1996. Acorn production by oaks in central coastal California: influence of weather at three levels. Canadian Journal of Forest Research 26: 1677-1683.

76. Koenig, W. D. and J. L. Dickinson.1996. Nestling sex-ratio variation in western bluebirds. Auk 113: 902-910.

77. Koenig, W. D., D. Van Vuren and P. N. Hooge. 1996. Detectability, philopatry, and the distribution of dispersal distances in vertebrates. Trends in Ecology & Evolution 11: 514-517. pdf

78. Lidicker, W. Z., Jr. and W. D. Koenig. 1996. Responses of terrestrial vertebrates to habitat edges and corridors. Pp. 85-109 in Metapopulations and Wildlife Conservation (D. R. McCullough, ed.) Island Press, Washington, D.C.

79. Koenig, W. D. 1997. Host preferences and behavior of oxpeckers: coexistence of similar species in a fragmented landscape. Evolutionary Ecology 11: 91-104. pdf

80. Monk, D. S. and W. D. Koenig. 1997. Individual, brood, and sex variation in begging calls of western bluebirds. Wilson Bulletin 109: 328-332.

81. Fincke, O. M., J. K. Waage and W. D. Koenig. 1997. Natural and sexual selection components of Odonate mating patterns. Pp. 58-74 in The Evolution of Mating Systems in Insects and Arachnids (J. C. Choe and B. J. Crespi, eds.) Cambridge University Press, Cambridge.

82. Knops, J. M. H., W. D. Koenig, and T. H. Nash, III. 1997. On the relationship between nutrient use efficiency and fertility in forest ecosystems. Oecologia 110: 550-556.

83. Knops, J. M. H. and W. D. Koenig. 1997. Site fertility and leaf nutrients of sympatric evergreen and deciduous species of Quercus in central coastal California. Plant Ecology 130: 121-131.

84. Reynolds, M. D. and W. D. Koenig. 1997. Reproductive success of yellow-billed magpies Pica nuttalli in central coastal California. Acta Ornithologica 32: 99-109.

85. Koenig, W. D. and J. M. H. Knops. 1997. Patterns of geographic synchrony in growth and reproduction of oaks within California and beyond. Pp.101-108 in Proceedings of the Symposium on Oak Woodlands: Ecology, Management, and Urban Interface Issues. (N. H. Pillsbury, J. Verner, and W. D. Tietje, Technical coordinators) Pacific SW For. & Range Exp. Stn Gen. Tech. Rep. PSW-GTR-160.

86. Koenig, W. D., J. Haydock, and M. T. Stanback. 1998. Reproductive roles in the cooperatively breeding acorn woodpecker: incest avoidance vs. reproductive competition. American Naturalist 151: 243-255. pdf

87. Koenig, W. D. and J. M. H. Knops. 1998. Testing for spatial autocorrelation in ecological studies. Ecography 21: 423-429. pdf

88. Koenig, W. D. 1998. Spatial autocorrelation in California land birds. Conservation Biology 12: 612-620

89. Koenig, W. D. and S. H. Faeth. 1998. Effects of storage on tannin and protein content of cached acorns. Southwestern Naturalist 43: 170-175.

90. Koenig, W. D. and J. M. H. Knops. 1998. Scale of mast-seeding and tree-ring growth. Nature 396: 225-226. pdf

91. Koenig, W. D. 1999. Spatial autocorrelation in ecological studies. Trends in Ecology & Evolution 14: 22-26. pdf

92. Hooge, P. N., M. T. Stanback and W. D. Koenig. 1999. Nest-site selection in the acorn woodpecker. Auk 116: 45-54.

93. Koenig, W. D. and J. R. Walters. 1999. Sex-ratio selection in species with helpers at the nest: the repayment model revisited. American Naturalist 153: 124-130.

94. Koenig, W. D. and J. Haydock. 1999. Oaks, acorns, and the geographical ecology of the acorn woodpecker. Journal of Biogeography 26: 159-165. pdf

95. Koenig, W. D., M. T. Stanback, and J. Haydock. 1999. Demographic consequences of incest avoidance in the cooperatively breeding acorn woodpecker. Animal Behaviour 57: 1287-1293.

96. Koenig, W. D., J. M. H. Knops, W. J. Carmen, and M. T. Stanback. 1999. Spatial dynamics in the absence of dispersal: acorn production by oaks in central coastal California. Ecography 22: 499-506.

97. Koenig, W. D., D. R. McCullough, C. E. Vaughn, J. M. H. Knops, and W. J. Carmen. 1999. Synchrony and asynchrony of acorn production at two coastal California sites. Madroño 46: 20-24.

98. Koenig, W. D. and J. M. H. Knops. 1999. The ecology of blue oak (Quercus douglasii) acorn production. Journal of the International Oak Society 9: 60-68.

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99. Koenig, W. D. and J. M. H. Knops. 2000. Patterns of annual seed production by northern hemisphere trees: a global perspective. American Naturalist 155: 59-69. pdf

Data from this paper: Site summary (pdf) ||| Data citations (pdf) | | | Data (.xls)

100. Koenig, W. D., P. N. Hooge, M. T. Stanback, and J. Haydock. 2000. Natal dispersal in the cooperatively breeding acorn woodpecker. Condor 102: 492-502.

101. Trenham, P. C., H. B. Shaffer, W. D. Koenig, and M. R. Stromberg. 2000. Life history and demographic variation in the California tiger salamander (Ambystoma californiense). Copeia 2000: 365-377. pdf

102. Knops, J. M. H. and W. D. Koenig. 2000. Annual variation in xylem water potential in California oaks. Madroño 47: 106-108.

103. Koenig, W. D., M. T. Stanback, J. Haydock, and F. Kraaijeveld-Smit. 2001. Nestling sex-ratio variation in the cooperatively breeding acorn woodpecker (Melanerpes formicivorus). Behavioral Ecology & Sociobiology 49: 357-365.

104. Haydock, J., W. D. Koenig, and M. T. Stanback. 2001. Shared parentage and incest avoidance in the cooperatively breeding acorn woodpecker. Molecular Ecology 10: 1515-1526.

105. Koenig, W. D. and J. M. H. Knops. 2001. Seed-crop size and eruptions of North American boreal seed-eating birds. Journal of Animal Ecology 70: 609-620.

106. Koenig, W. D. 2001. Synchrony and periodicity of eruptions by boreal seed-eating birds. Condor 103: 725-735.

107. Koenig, W. D. 2001. Spatial autocorrelation and local disappearances in wintering North American birds. Ecology 82: 2636-2644. pdf

108. Trenham, P. C., W. D. Koenig, and H. B. Shaffer. 2001. Spatially autocorrelated demography and interpond dispersal in the pond breeding salamander Ambystoma californiense. Ecology 82: 3519-3530. pdf

109. Koenig, W. D. and J. M. H. Knops. 2002. The behavioral ecology of masting in oaks. Pp. 129- 148 in Oak Forest Ecosystems (W. J. McShea and W. M. Healy, eds.) The Johns Hopkins University Press, Baltimore, MD.

110. Koenig, W. D. 2002. Global patterns of environmental synchrony and the Moran effect. Ecography 25: 283-288.pdf

111. Koenig, W. D., J. M. H. Knops, and W. J. Carmen. 2002. Arboreal seed removal and insect damage in three California oaks. Pp. 193-204 in Proceedings of the Fifth Symposium on Oak Woodlands: Oaks in California’s Changing Landscape. R. B. Standiford, D. McCreary, and K. L. Purcell, technical coordinators. Pacific SW Forest & Range Exp. Station Gen. Tech. Rep. PSW-GTR-184.

112. Koenig, W. D. and L. S. Benedict. 2002. Size, insect parasitism, and energetic value of acorns stored by acorn woodpeckers. Condor 104: 539-547.

113. Chu, M., W. D. Koenig, A. Godinez, C. E. McIntosh, and R. C. Fleischer. 2002. Social and genetic monogamy in territorial and loosely colonial populations of the phainopepla. Auk  119: 770-777.

114. Haydock, J. and W. D. Koenig. 2002. Reproductive skew in the polygynandrous acorn woodpecker. Proceedings of the National Academy of Sciences (USA) 99: 7178-7183.pdf

115. Craft, K. J., M. V. Ashley, and W. D. Koenig. 2002. Limited hybridization between Quercus lobata and Quercus douglasii (Fagaceae) in a mixed stand in central coastal California. American Journal of Botany 89: 1792-1798.pdf

116. Koenig, W. D. and M. V. Ashley. 2003. Is pollen limited? The answer is blowin’ in the wind. Trends in Ecology & Evolution 18: 157-159.pdf

117. Haydock, J. and W. D. Koenig. 2003. Patterns of reproductive skew in the polygynandrous acorn woodpecker. American Naturalist 162: 277-289.pdf

118. Trenham, P. C., W. D. Koenig, M. J. Mossman, S. L. Stark, and L. A. Jagger. 2003. Regional dynamics of wetland-breeding frogs and toads: turnover and synchrony. Ecological Applications 13: 1522-1532. pdf

119. Koenig, W. D. and A. M. Liebhold. 2003. Regional impacts of periodical cicadas on oak radial increment. Canadian Journal of Forest Research 33: 1084-1089. pdf

120. Koenig, W. D. 2003. European starlings and their effect on native cavity-nesting birds. Conservation Biology 17: 1134-1140. pdf

121. Koenig, W. D., D. Kelly, V. L. Sork, R. P. Duncan, J. S. Elkinton, M. S. Peltonen and R. D. Westfall. 2003. Dissecting components of population-level variation in seed production and the evolution of masting behavior. Oikos 102: 581-591. pdf

122. Buonaccorsi, J. P., J. Elkinton, W. Koenig, R. Duncan, D. Kelly and V. Sork. 2003. Measuring mast seeding behavior: relationships among population variation, individual variation and synchrony. Journal of Theoretical Biology 224: 107-114.

123. Koenig, W. D. and J. L. Dickinson. 2004. Introduction. Pp. 1-4 in Ecology and Evolution of Cooperative Breeding in Birds (W. D. Koenig and J. L. Dickinson, eds.). Cambridge University Press, Cambridge.

124. Koenig, W. D. and J. Haydock. 2004. Incest avoidance. Pp. 142-156 in Ecology and Evolution of Cooperative Breeding in Birds (W. D. Koenig and J. L. Dickinson, eds.). Cambridge University Press, Cambridge.

125. Liebhold, A. M., V. Sork, M. Peltonen, W. D. Koenig, O. Bjørnstad, R.  Westfall, J.  Elkinton, and J. M. H. Knops. 2004. Within-population spatial synchrony in mast seeding of North American oaks. Oikos 104: 156-164. pdf

126. Liebhold, A. M., W. D. Koenig and O. Bjørnstad. 2004. Spatial synchrony in population dynamics. Annual Review of Ecology, Evolution, and Systematics 35: 467-490. pdf

127. Koenig, W. D. and A. M. Liebhold. 2005. Effects of periodical cicada emergences on abundance and synchrony of avian populations. Ecology 86: 1873-1882. pdf

128. Koenig, W. D., E. L. Walters, J. R. Walters, J. S. Kellam, K. G. Michalek, and M. S. Schrader. 2005. Seasonal body weight variation in five species of woodpeckers. Condor 107: 810-822. pdf

129. Monahan, W. B. and W. D. Koenig. 2006. Potential effects of SOD on the oak woodland bird community of coastal California. Pp. 195-209 in Proceedings of the Sudden Oak Death Second Science Symposium: the State of Our Knowledge. (S. J. Frankel, P. J. Shea, and M. I. Haverty, Technical coordinators). Pacific SW Forest & Range Exp. Station Gen. Tech. Rep. PSW-GTR-196.

130. Monahan, W. B. and W. D. Koenig. 2006. Estimating the effects of sudden oak death on oak-dependent birds. Biological Conservation 127: 146-157. pdf

131. Koenig, W. D., M. T. Stanback, and J. Haydock. 2006. Abbreviated inner primaries: a sex-linked dimorphism in the acorn woodpecker. Journal of Field Ornithology 77: 157-162. pdf

132. Koenig, W. D. 2006. Spatial synchrony of monarch butterflies. American Midland Naturalist 155: 39-49. pdf

133. Wiebe, K. L., W. D. Koenig, and K. Martin. 2006. Evolution of clutch size in cavity-excavating birds: the nest site limitation hypothesis revisited. American Naturalist 167: 343-353. pdf

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134. Koenig, W. D., L. Marcus, T. W. Scott, and J. L. Dickinson. 2007. West Nile virus and California breeding bird declines. EcoHealth 4: 18-24. pdf

135. Koenig, W. D. and J. T. Stahl. 2007. Late summer and fall nesting in the acorn woodpecker and other North American terrestrial birds. Condor 109: 334-350.pdf

136. Wiebe, K., W. D. Koenig, and K. Martin. 2007. Costs and benefits of nest reuse versus excavation in cavity-nesting birds. Annales Zoologici Fennici 44: 209-217.pdf

137. Knops, J. M. H., W. D. Koenig, and W. J. Carmen. 2007. A negative correlation does not imply a trade-off between growth and reproduction in California oaks. Proceedings of the National Academy of Sciences (USA) 104: 16982-16985. pdf

138. Koenig, W. D. and J. M. H. Knops. 2007. Long-term growth and persistence of blue oak (Quercus douglasii) seedlings in a California oak savanna. Madroño 54: 269-274. pdf

139. Koenig, W. D. and J. M. H. Knops. 2008. Timing of flowering and seed production in three California oaks. Pp. 371-380 in A. Merenlender, D. McCreary, and K. L. Purcell (tech. coords.),  Proceedings of the Sixth Symposium on Oak Woodlands: Today’s Challenges, Tomorrow’s Opportunities. Pacific SW Forest & Range Exp. Station Gen. Tech. Rep. PSW-GTR-217.

140. Garrison, B. A., W. D. Koenig, and J. M. H. Knops. 2007. Spatial synchrony and temporal patterns in acorn production by California black oaks. Pp. 343-356 in A. Merenlender, D. McCreary, and K. L. Purcell (tech. coords.), Proceedings of the Sixth Symposium on Oak Woodlands: Today’s Challenges, Tomorrow’s Opportunities. Pacific SW Forest & Range Exp. Station Gen. Tech. Rep. PSW-GTR-217.

141. Scott, T., P.-Y. Lee, K. Paggett, R. Carney, S. Husted, and W. D. Koenig. 2007. The impact of West Nile virus on birds in California’s hardwood rangelands. Pp. 151-163 in Proceedings of the Sixth Symposium on Oak Woodlands: Today’s Challenges, Tomorrow’s Opportunities. Pacific SW Forest & Range Exp. Station Gen. Tech. Rep. PSW-GTR-217.

142. Koenig, W. D., and J. L. Dickinson. 2008. Cooperative breeding as an alternative reproductive tactic. Pp. 451-470 in Alternative Reproductive Tactics: an Integrative Approach (R. Oliveira, M. Taborsky, and J. Brockmann, eds.). Cambridge University Press, Cambridge.

143. Koenig, W. D., S.-F. Shen, A. H. Krakauer, and J. Haydock. 2009. Reproductive skew in avian societies. Pp. 227-264 in Reproductive Skew in Vertebrates: Proximate and Ultimate Causes (R. Hager and C. B. Jones, eds.). Cambridge University Press, Cambridge.

144. Crosbie, S. P., W. D. Koenig, K. Reisen, V. L. Kramer, L. Marcus, R. Carney, E. Pandolfino, G. M. Bolen, L. R. Crosbie, D. A. Bell, H. B. Ernest. 2008. Early impact of West Nile virus on the yellow-billed magpie (Pica nuttalli). Auk 125: 542-550.

145. Stromberg, M. R., W. D. Koenig, E. L. Walters, and J. Schweisinger. 2008. Estimate of Trichomonas gallinae-induced mortality in band-tailed pigeons, upper Carmel Valley, California, winter 2006-2007. Wilson Journal of Ornithology 120: 603-606.

146. Koenig, W. D. 2008. Lifetime reproductive success and sexual selection theory. Pp. 153-166 in Dragonflies and damselflies: model organisms for ecological and evolutionary research (A. Córdoba-Aguilar, ed.) Oxford University Press, Oxford, UK.

147. Koenig, W. D., D. J. Shaefer, S. Mambelli, and T. E. Dawson. 2008. Acorns, insects, and the diet of adult vs. nestling acorn woodpeckers. Journal of Field Ornithology79: 280-285.

148. Monahan, W. B., J. Tse, W. D. Koenig, and M. Garbelotto. 2008. Herbarium specimens suggest non-native origins for three species of Phytophthora in California. Mycological Research112: 757-758.

149. Koenig, W. D., J. P. McEntee, and E. L. Walters. 2008. Acorn harvesting by acorn woodpeckers: annual variation and comparison with genetic estimates. Evolutionary Ecology Research 10: 811-822. pdf

150. Kelly, D., W. D. Koenig, and A. M. Liebhold. 2008. An intercontinental comparison of the dynamic behavior of mast seeding communities. Population Ecology 50: 329-342. pdf

151. Koenig, W. D., A. H. Krakauer, W. B. Monahan, J. Haydock, J. M. H. Knops, and W. J. Carmen. 2009. Mast-producing trees and the geographical ecology of western scrub-jays. Ecography 32: 561-570. pdf

152. Koenig, W. D., J. M. H. Knops, W. J. Carmen, and R. D. Sage. 2009. No trade-off between seed size and number in the valley oak Quercus lobata. American Naturalist173: 682-688.pdf

153. Koenig, W. D., J. M. H. Knops, J. L. Dickinson, and B. Zuckerberg. 2009. Latitudinal decrease in acorn size in bur oak (Quercus macrocarpa) is due to environmental constraints, not avian dispersal. Botany 87: 349-356.pdf

154. Koenig, W. D., E. L. Walters, and J. Haydock. 2009. Helpers and egg investment in the cooperatively breeding acorn woodpecker: testing the concealed helper effects hypothesis. Behavioral Ecology and Sociobiology 63: 1659-1665.pdf

155. Koenig, W. D., and M. D. Reynolds. 2009. Yellow-billed Magpie (Pica nuttalli), in The Birds of North America Online (A. Poole, Ed.). Cornell Lab of Ornithology, Ithaca, NY. http://bna.birds.cornell.edu/bna/species/180 (doi:10.2173/bna.180)

156. Koenig, W. D., W. M. Hochachka, B. Zuckerberg, and J. L. Dickinson. 2010. Ecological determinants of American crow mortality due to West Nile virus during its North American sweep. Oecologia 163: 903-909. pdf

157. Koenig, W. D., J. M. H. Knops and W. J. Carmen. 2010. Testing the environmental prediction hypothesis for mast-seeding in CAlifornia oaks. Canadian Journal of Forest Research 40: 2115-2122 pdf

158. Koenig, W. D., L. Ries, V. B. K. Olsen, and A. M. Liebhold. 2011. Avian predators are less abundant during periodicial cicada emergences, but why? Ecology 92: 784-790. pdf

159. Sage, R. D., W. D. Koenig, and B. C. McLaughlin. 2011. Fitness consequences of seed size in the valley oak Quercus lobata Née (Fagaceae). Annals of Forest Science 68: 477-484.pdf

160. Nunn, C. L., V. O. Ezenwa, C. Arnold, and W. D. Koenig. 2011. Mutualism or parasitism? Using a phylogenetic approach to characterize the oxpecker-ungulate relationship. Evolution 65: 1297-1304.pdf

161. Zuckerberg, B., D. N. Bonter, W. M. Hochachka, W. D. Koenig, A. T. DeGeatano, and J. L. Dickinson. 2011. Climatic constraints on wintering bird distributions are modified by urbanization and weather. Journal of Animal Ecology 80: 403-413.pdf

162. Scofield, D. G., V. R. Alfaro, V. L. Sork, D. Grivet, E. Martinez, J. Papp, A. R. Pluess, W. D. Koenig, and P. E. Smouse. 2011. Foraging patterns of acorn woodpeckers (Melanerpes formicivorus) on valley oak (Quercus lobata Née) in two California oak savanna-woodlands. Oecologia 166: 187-196.pdf

163. Rosenstock, T. S., A. Hastings, W. D. Koenig, D. J. Lyles, and P. H. Brown. 2011. Testing Moran’s theorem in an agroecosystem. Oikos 120: 1434-1440.pdf

164. Abraham, S. T., D. N. Zaya, W. D. Koenig, and M. V. Ashley. 2011. Interspecific and intraspecific pollination patterns of valley oak, Quercus lobata, in a mixed stand in coastal central California. International Journal of Plant Sciences 172: 691-699.pdf

165. Koenig, W. D., E. L. Walters, and A. M. Liebhold. 2011. Effects of gypsy moth outbreaks on North American woodpeckers. Condor 113: 352-361. pdf

166. Koenig, W. D., E. L. Walters, and J. Haydock. 2011. Variable helper effects, ecological conditions, and the evolution of cooperative breeding in the acorn woodpecker. American Naturalist 178: 145-158. pdf

167. Morrison, S. A., T. S. Sillett, C. K. Ghalambor, J. W. Fitzpatrick, D. M. Graber, V. J. Bakker, R. Bowman, C. T. Collins, P. W. Collins, K. S. Delaney, D. F. Doak, W. D. Koenig, L. Laughrin, A. A. Lieberman, J. M. Marzluff, M. D. Reynolds, J. M. Scott, J. A. Stallcup, W. Vickers, and W. M. Boyce. 2011. Proactive conservation management of an island-endemic bird species in the face of global change. BioScience 61: 1013–1021.pdf

168. Koenig, W. D., and E. L. Walters. 2011. Age-related provisioning behaviour in the cooperatively breeding acorn woodpecker: testing the skills and the pay-to-stay hypotheses. Animal Behaviour 82: 437-444. pdf

169. Koenig, W. D., E. L. Walters, and J. Haydock. 2011. Fitness consequences of within-brood dominance in the cooperatively breeding acorn woodpecker. Behavioral Ecology and Sociobiology 65: 2229-2238. pdf

170. Koenig, W. D., and E. L. Walters. 2012. Brooding, provisioning, and compensatory care in the cooperatively breeding acorn woodpecker. Behavioral Ecology 23: 181-190. pdf

171. Zuckerberg, B., A. Desrochers, W. M. Hochachka, W. D. Koenig, and J. L. Dickinson. 2012. Do overlapping landscapes lead to a violation in spatial dependency? Journal of Wildlife Management 76: 1072-1080.pdf

172. Koenig, W. D., K. A. Funk, T. S. Kraft, W. J. Carmen, B. C. Barringer, and J. M. H. Knops. 2012. Stabilizing selection for within-season flowering phenology confirms pollen limitation in a wind-pollinated tree. Journal of Ecology 100: 758-763. pdf

173. Koenig, W. D., and E. L. Walters. 2012. An experimental study of chick provisioning in the cooperatively breeding acorn woodpecker. Ethology 118: 566-574. pdf

174. Barringer, B. C., W. D. Koenig, and J. M. H. Knops. 2012. Interrelationships among life-history traits in three California oaks. Oecologia 171: 129-139. pdf

175. Knops, J.M. H. and W. D. Koenig. 2012. Sex allocation in California oaks: trade-offs or resource tracking? PLoS One 7(8): e43492. pdf

176. Koenig, W. D. and A. M. Liebhold. 2013. Avian predation pressure as a potential driver of periodical cicada cycle length. American Naturalist 181: 145-149. pdf

177. Koenig, W. D. and J. M. H. Knops. 2013. Large scale spatial synchrony and cross-synchrony in acorn production by two California oaks. Ecology 94: 83-93. pdf

178. Rusk, C. L., E. L. Walters, and W. D. Koenig. 2013. Cooperative breeding and long-distance dispersal: a test using vagrant records. PLoS One 8(3): e58624. doi:10.1371/journal.pone.0058624 pdf

179. Koenig, W. D., A. M. Liebhold, D. N. Bonter, W. M. Hochachka, and J. L. Dickinson. 2013. Effects of the emerald ash borer invasion on four species of birds. Biological Invasions 15: 2095-2103. doi:10.1007/s10530-013-0435-x pdf

180. Pearse, I. S., S. Griswold, D. Pizarro, and W. D. Koenig. 2014. Stage and size structure of three species of oaks in central coastal California. Madroño 61: 1-8. pdf Data from this paper are available here.

181. Pearse, I. S., W. D. Koenig, and J. M. H. Knops. 2014. Cues versus proximate drivers: testing the mechanism behind masting behavior. Oikos 123: 179-184. pdf

182. W. D. Koenig, M. Díaz, F. Pulido, R. Alejano, E. Beamonte, and J. M. H. Knops. 2013. Acorn production patterns. Pp. 181-209 in Mediterranean Oak Woodland Working Landscapes: Dehesas of Spain and Ranchlands of California, P. Campos, L. Huntsinger, J. L. Oviedo, P. F. Starrs, M. Díaz, R. B. Standiford, and G. Montero, editors. Springer Landscape Series, vol. 16. pdf

183. Koenig, W. D. and J. M. H. Knops. 2014. Environmental correlates of acorn production by four species of Minnesota oaks. Population Ecology 56: 63-71. pdf

184. Koenig, W. D., E. L. Walters, I. S. Pearse, W. J. Carmen, and J. M. H. Knops. 2014. Serotiny in California oaks. Madroño 61: 151-158. pdf

185. Koenig, W. D. and E. L. Walters. 2014. What we don't know, and what needs to be known, about the cooperatively breeding acorn woodpecker. Acta Ornithologica 49: 221-232. pdf

186. Koenig, W. D., J. M. H. Knops, W. J. Carmen, and I. S. Pearse. 2015. What drives masting? The phenological synchrony hypothesis. Ecology 96: 184-192. pdf

187. Pearse, I. S., W. D. Koenig, K. A. Funk, and M. A. Pesendorfer. 2015. Pollen limitation and flower abortion in a wind-pollinated, masting tree. Ecology 96: 587-593. pdf

188. Pearse, I. S., K. A. Funk, and T. S. Kraft, and W. D. Koenig. 2015. Lagged effects of early season herbivores on valley oak fecundity. Oecologia 178: 361-368. pdf

189. Koenig, W. D., E. L. Walters, J. M. H. Knops, and W. J. Carmen. 2015. Acorns and acorn woodpeckers: ups and downs in a long-term relationship. In: Proceedings and the 7th California Oak Symposium: Managing Oak Woodlands in a Dynamic World. Pacific SW Forest & Range Exp. Station Gen. Tech. Rep.

190. Pérez-Ramos, I. M., C. M. Padilla-Díaz, W. D. Koenig, and T. Marañón. 2015. Environmental drivers of mast-seeding in Mediterranean oak species: does leaf habit matter? Journal of Ecology 103: 691-700. pdf

191. Strong, C., B. Zuckerberg, J. L. Betancourt, and W. D. Koenig. 2015. Climatic dipoles driver two principal modes of North American boreal bird irruption. Proceedings of the National Academy of Sciences (USA) 112: E2795-E2802. pdf

192. McMahon, D. E., I. S. Pearse, W. D. Koenig, and E. L. Walters. 2015. Tree community shifts and acorn woopecker population increases over three decades in a California oak woodland. Canadian Journal of Forest Research 45:1113-1120. pdf

193. Pearse, I. S., J. H. Baty, D. Herrmann, R. Sage, and W. D. Koenig. 2015. Leaf phenology mediates provenance differences in herbivore populations on valley oaks in a common garden. Ecological Entomology 40: 525-531. pdf

194. Koenig, W. D. and E. L. Walters. 2015. Temporal variability and cooperative breeding: testing the bet-hedging hypothesis in the acorn woodpecker. Proceedings of the Royal Society B 282: 20151742. pdf

195. Ashley, M. V., S. T. Abraham, J. R. Backs, and W. D. Koenig. 2015. Landscape genetics and population structure in valley oak (Quercus lobata Née). American Journal of Botany 102: 21243-2131. pdf

196. Funk, K. A., W. D. Koenig, and J. M. H. Knops. 2016. Fire effects on acorn production are consistent with the stored resource hypothesis for masting behavior. Canadian Journal of Forest Research 46: 20-24. pdf

197. Dickinson, J. L. and W. D. Koenig. 2016. Introduction. Pp. 1-5 in Cooperative Breeding in Vertebrates: Studies in Ecology, Evolution, and Behavior. (W. D. Koenig and J. L. Dickinson, eds.). Cambridge University Press, Cambridge.

198. Koenig, W. D., E. L. Walters and J. Haydock. 2016. Acorn woodpeckers: helping at the nest, polygynandry, and dependence on a variable acorn crop. Pp. 217-236 in Cooperative Breeding in Vertebrates: Studies in Ecology, Evolution, and Behavior. (W. D. Koenig and J. L. Dickinson, eds.). Cambridge University Press, Cambridge. pdf

199. Koenig, W. D., J. L. Dickinson and S. T. Emlen. 2016. Synthesis: cooperative breeding in the 21st century. Pp. 353-374 in Cooperative Breeding in Vertebrates: Studies in Ecology, Evolution, and Behavior. (W. D. Koenig and J. L. Dickinson, eds.). Cambridge University Press, Cambridge. pdf

200. Pesendorfer, M. B., T. S. Sillett, W. D. Koenig and S. A. Morrison. 2016. Scatter-hoarding corvids as seed dispersers for oaks and pines: a review of a widely distributed mutualism and its utility to habitat restoration. Condor: Ornithological Applications 118: 215-237. pdf

201. Rogers, K. H., Y. A. Girard, W. D. Koenig, and C. K. Johnson. 2016. Ecological drivers and population impacts of avian Trichoonosis mortality events in band-tailed pigeons (Patagioenas monilis) in California, USA. Journal of Wildlife Diseases 52: 484-494. pdf

202. Koenig, W. D. and A. M. Liebhold. 2016. Temporally increasing spatial synchrony of North American temperature and bird populations. Nature Climate Change 6: 614-617. pdf

203. Pesendorfer, M. B., W. D. Koenig, I. S. Pearse, J. M. H. Knops, and K. A. Funk. 2016. Individual resource-limitation combined with population-wide pollen availability drives masting in the valley oak (Quercus lobata). Journal of Ecology 104: 637-645. pdf

204. Pesendorfer, M. B. and W. D. Koenig. 2016. The effect of within-year variation in acorn crop size on seed harvesting by avian horders. Oecologia 181: 97-106. pdf

205. Koenig, W. D. and E. L. Walters. 2016. Provisioning patterns in the cooperatively-breeding acorn woodpecker: is feeding a signal, and if so, to whom and why? Animal Behaviour 119: 125-134. pdf

206. Koenig, W. D., R. Alejano, M. D. Carbonero, P. Fernández-Rebollo, J. M. H. Knops, T. Marañón, C. M. Padilla-Díaz, I. S. Pearse, I. M. Pérez-Ramos, J. Vázquez-Piqué, and M. B. Pesendorfer. 2016. Is the relationship between mast-seeding and weather in oaks related to their life-history or phylogeny? Ecology 97: 2603-2615. pdf

207. Pearse, I. S., W. D. Koenig, and D. Kelly. 2016. [Tansley review] Mechanisms of mast seeding: resources, weather, cues, and selection. New Phytologist 212: 546-562. pdf

208. Pesendorfer, M. B., and W. D. Koenig, 2017. Competing for seed dispersal: evidence for the role of avian seed hoarders in mediating apparent predation among oaks. Functional Ecology 31: 622-631. pdf

209. Koenig, W. D. J. M. H. Knops, W. J. Carmen, and M. B. Pesendorfer. 2017. Testing the terminal investment hypothesis in California oaks. American Naturalist 189: 564-569. pdf

210. Koenig, W. D., E. L. Walters and P. G. Rodewald. 2017. Testing alternative hypotheses for the cause of population declines: the case of the red-headed woodpecker. Condor: Ornithological Applications 119: 143-154. pdf

211. Gharehaghaji, M., E. S. MInor, M. V. Ashley S. T. Abraham, and W. D. Koenig, 2017. Effects of landscape features on gene flow of valley oaks (Quercus lobata). Plant Ecology 218: 487-499. pdf

212. Koenig, W. D. and A. M. Liebhold. 2017. A decade of emerald ash borer effects on regional woodpecker and nuthatch populations. Biological Invasions 19: 2029-2037. pdf

213. Cockburn, A., B. J. Hatchwell, and W. D. Koenig, 2016. Sociality in birds.Pp. 320-353 in Comparative Social Evolution. (D. R. Rubenstein and P. Abbot, eds.). Cambridge University Press, Cambridge.

214. Shen, S.-F., S. T. Emlen, W. D. Koenig, and D. R. Rubenstein. 2017. The ecology of cooperative breeding behavior. Ecology Letters 20: 708-720. pdf

215. Koenig,W. D. 2017. Striving for science that is transparent, credible--and enjoyable: a comment on Ihle et al. Behavioral Ecology 28: 358. pdf

216. Koenig, W. D., J. M. H. Knops, M. B. Pesendorfer, D. N. Zaya, and M. V. Ashley. 2017. Drivers of synchrony of acorn production in the valley oak (Quercus lobata) at two spatial scales. Ecology 98: 3056-3062. pdf

217. Pearse, I. S., J. M. LaMontagne, and W. D. Koenig. 2017. Interannual variation in seed production has increased over time. Proceedings of the Royal Society B 284: 20171666. doi: 10.1098/rspb.2017.1666. pdf

218. Pesendorfer, M. B. and W. D. Koenig. J. 2018. Does aggression avoidance drive oak tree attendance by corvid scatter-hoarders? Behavioral Ecology and Sociobiology 72: 85. doi: 10.1007/s00265-018-2498-2. pdf

219. Koenig, W. D., J, J. M. H. Knops, W. J. Carmen, M. B. Pesendorfer, and J. L. Dickinson. 2018. Effects of mistletoe (Phoradendron villosum) on California oaks. Biology Letters 14: 20180240. pdf

220. Pardo, M.A., E. A. Sparks, T. S. Kuray, N. D. Hagemeyer, E. L. Walters, and W. D. Koenig. J. 2018. Wild acorn woodpeckers recognize associations between individuals in other groups. Proceedings of the Royal Society B 285: 20181017. doi: 10.1098/rspb.2018.1017 pdf

221. Schubert, S. C., M. B. Pesendorfer, and W. D. Koenig. 2018. Context-dependent post-dispersal predation of acorns in a California oak community. Acta Oecologica 92: 52-58. pdf

222. Koenig, W. D. and E. L. Walters. 2018. Causes of seasonal decline in reproduction of the cooperatively-breeding acorn woodpecker. Journal of Avian Biology 49: e01784. pdf

223. Moreria, X., L. Abdala-Roberts, I. M. Pérez-Ramos, J. M. H. Knops, M. B. Pesendorfer, W. D. Koenig , and K. A. Mooney. 2019. Weather cues associated with masting behavior dampen the negative autocorrelation between past and current reproduction in oaks. American Journal of Botany 106: 51-60. pdf

224. Pesendorfer, M. B., M. Bogdziewicz, W. D. Koenig, M. Ledwon and M. Zywiec. 2019. Declining fruit production before death in a widely distributed tree. Annals of Forest Science 76:11. pdf

225. Barve, S., W. D. Koenig, J. Haydock and E. L. Walters. 2019. Habitat saturation results in joint-nesting female coalitions in a social bird. American Naturalist 193: 830-840. pdf

226. Koenig, W. D., E. L. Walters, and S. Barve. 2019. Does helping-at-the-nest help? The case of the acorn woodpecker. Frontiers in Ecology and Evolution 7:272. pdf

227. Koenig, W. D., E. L. Walters, P. B.Stacey, M. T. Stanback, and R. L. Mumme. 2019. Acorn Woodpecker (Melanerpes formicivorus), version 2.0. In: Birds of North America (P. G. Rodewald, Editor) Cornell Lab of Ornithology, Ithaca, NY. https://doi-org.proxy.library.cornell.edu/10.2173/bna.acowoo.02

228. Fernández-Martínez, M., I. Pearse, I., J. Sardans, F. Sayol, W. D. Koenig, J. M. LaMontagne, M. Bogdziewicz, A. Collalti, A. Hacket-Pain, G. Vacchiano, J. M. Espelta, J. Peñuelas, and I. A. Janssens. 2019. Nutrient scarcity as a selective pressure for mast seeding. Nature Plants  5: 1222-1228. pdf

229. Barve, S., N. D. G. Hagemeyer, R. E. Winter, S. D. Chamberlain, W. D. Koenig, and E. L. Walters. 2020. Wandering woodpeckers: foray behavior in a social bird. Ecology  101: e02943. pdf

230. Barringer, B. C., W. D. Koenig, , I. S. Pearse, and J. M. H. Knops. 2020. Population ecology and spatial synchrony in the abundance of leaf gall wasps within and among populations of valley oak. (Quercus lobata). Population Ecology  62: 220-232. pdf

231. Bogdziewicz, M., D. Ascoli, A. Hacket-Pain, W. D. Koenig, M. B. Pesendorfer, I. S. Pearse, A. Satake, P. Thomas, G. Vacchiano, T. Wohlgemuth, and A. Tanentzap. 2020. From theory to experiments for testing the proximate mechanisms of mast seeding: an agenda for an experimental ecology. Ecology Letters  23: 210-220. pdf

232. Zuckerberg, B., C. Strong, J. M. LaMontagne, S. St. George, J. L. Betancourt, and W. D. Koenig. 2020. Climate dipoles as continental drivers of plant and animal populations. Trends in Ecology & Evolution   35: 440-453. pdf

233. LaMontagne, J., I. S. Pearse, D. F. Greene, and W. D. Koenig. . 2020. Mast seeding patterns are asynchronous at a continental scale. Nature Plants  6: 460-465. pdf

234. Pearse, I.S., J. M. LaMontagne, M. Lordon, A. L. Hipp, and W. D. Koenig. 2020. Biogeography and phylogeny of masting: do global patterns fit functional hypotheses? New Phytologist  227: 1557-1567. doi: 10.1111/nph.16617. pdf

235. Koenig, W. D., J. M. H. Knops, and W. J. Carmen 2020. Can mast history be inferred from radial growth? A test using five species of California oaks. Forest Ecology and Management  472: 118233. doi: 10.1016/j.foreco.2020.118233. pdf

236. Pardo, M. A., C. E. Hayes, E. L. Walters, and W. D. Koenig. 2020. Acorn woodpeckers vocally discriminate current and former group members from non-group members. Behavioral Ecology  31: 1120-1128. pdf

237. Koenig, W. D., J. M. H. Knops, and W. J. Carmen. 2020. Intraspecific variation in the relationship between weather and masting behavior in valley oak Quercus lobata. Canadian Journal of Forest Research 50: 1299-1306. pdf

238. Barve, S., A. S. Lahey, R. M. Brunner, W. D. Koenig, and E. L. Walters. 2020. Woodpecker wars: tracking warriors and spectators with telemetry. Current Biology  30: R982-R983. pdf

239. Pardo, M. A., E. L. Walters, and W. D. Koenig. 2020. Experimental evidence that acorn woodpeckers recognize relationships among third parties no longer living. Behavioral Ecology  31: 1257-1265. pdf

240. Koenig, W. D. 2020. What are the competitive effects of invasive species? Forty years of the Eurasian Collared-dove in North America. Biological Invasions  22: 3645-3652. pdf

241. Koenig, W. D. , E. M. Hallock, D. J. Weber, and E. L. Walters. 2021. Nest cavity reuse by the cooperatively breeding acorn woodpecker. Ornithology 138: ukaa088. pdf

242. Clark, J. S., et al. 2021. Continent-wide tree fecundity driven by indirect climate effects. Nature Communications  12:1242. pdf

243. Koenig, W. D. , A. C. B. Prinz, J. Haydock, H. L. Dugdale, and E. L. Walters. 2021. Are you my baby? Testing whether paternity affects behavior of cobreeder male acorn woodpeckers. Behavioral Ecology 32: 865-874. pdf

244. Koenig, W. D. 2021. A brief history of masting research. Philosophical Transactions of the Royal Society B376: 20200423. pdf

245. Koenig, W. D. , M. B. Pesendorfer, I. S. Pearse, and J. M. H. Knops. 2021. Budburst timing of valley oaks at Hastings Reservation, central coastal California. Madroño  68:434-442. pdf

246. Barve, S., C. Riehl, E. L. Walters, J. Haydock, H. L. Dugdate, and W. D. Koenig. 2021. Lifetime reproductive benefits of cooperative polygamy vary for males and females in the acorn woodpecer (Melanerpes formicivorus). Proceedings of the Royal Society B 288: 20210579. pdf

247. Qiu, T., et al. 2021. Is there tree senescence? The fecundity evidence. Proceedings of the National Academy of Science (USA) 118: e2106130118. pdf

248. Sharma, S., et al. 2022. North American tree migration paced by climate in the West, lagging in the East. Proceedings of the National Academy of Science (USA) 119: e216691118. pdf

249. Hacket-Pain, A., J. J. Foest, I. S. Pearse, J. M. LaMontagne, W. D. Koenig, et al. 2022. MASTREE+: time-series of plant reproductive effort from six continents. Global Change Biology 28: 3066-3082. pdf

250. Justino, M. A., N. D. G. Hagemeyer, M. T. Banik, J. M. Palmer, D. L. Lindner, M. E. Smith, W. D. Koenig, and E. L. Walters. 2022. Fungal communities associated with acorn woodpeckers and their excavations. Fungal Ecology 59: 101154 pdf

251. Koenig, W. D. and J. M. H. Knops. 2022. Drivers of winter population cycles in the varied thrush Ixoreus naevius. Canadian Journal of Zoology 100: 607-613. pdf

252. Koenig, W. D., A. M. Liebhold, J. M. LaMontagne, and I. S. Pearse. 2022. Periodical cicada emergences affect masting behavior of oaks. American Naturalist 201: 755-762. pdf

253. Koenig, W. D., S. Barve, J. Haydock, H. L. Dugdale, M. K. Oli, and E. L. Walters. 2023. Lifetime inclusive fitness effects of cooperative polygamy in the acorn woodpecker. Proceedings of the National Academy of Science (USA) 120: e2219345120. pdf

254. Halupka, L., et al. 2023. The effect of climate change on avian offspring: a global meta-analysis. Proceedings of the National Academy of Science (USA) 120: e2208389120. pdf

255. Hagemeyer, N.D.G., M. B. Pesendorfer, W. D. Koenig, and E. L. Walters 2023. Investigating the drivers of temporal occupancy-abundance relationships in a cooperatively breeding bird. Landscape Ecology   38: 1955-1970. pdf

256. Koenig, W. D., J. Haydock, H. L. Dugdale, and E. L. Walters 2023. Territory inheritance and the evolution of cooperative breeding in the acorn woodpecker. Animal Behaviour   205: 241-249. pdf

257. Huffmyer, W. L., F. Ji, J. C. Blackwood, A. Hstings, W. D. Koenig, A. M. Liebhold, M. Machta, and K. C. Abbott. 2023. Variation in avian predation pressure as a driver for the diversificatino of periodical cicada broods. American Naturalist   203: E92-E106. pdf

258. Nigro, K. M. et al. 2024.coMAST: harmonized seed production data for woody plants across U.S. long-term research sites. Ecology   (in press) pdf

259. LaMontagne, J., et al. 20204. Community synchrony in seed production is associated with trait similarity and climate across North America. Ecology Letters   (in press) pdf

260. Hagemeyer, N. D. G., W. D. Koenig, and E. L. Walters 2024. Active facilitation of helper dispersal by parents and siblings in the cooperatively breeding acorn woodpecker. Behavioral Ecology and Sociobiology   (in press) pdf

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News and Views-type articles

 

1. Koenig, W. D. and J. Haydock. 2001. Dividing up the kids. Science 291: 442-443.

2. Dickinson, J. L. and W. D. Koenig. 2003. Desperately seeking similarity. Science 300: 1887-1889.

3. Koenig, W. D. 2005. Persistence in adversity: lessons from the ivory-billed woodpecker. BioScience 55: 646-647.

4. Koenig, W. D. and E. L. Walters. 2008. A tale of two worlds: molecular ecology and population structure of the threatened Florida scrub-jay. Molecular Ecology 17: 1632-1633.

5. Koenig, W. D. 2017. What drives cooperative breeding? PLoS Biology 15(6): e2002965. pdf

General articles

 

  1. Stacey, P. B., and W. D. Koenig. 1984. Cooperative breeding in the acorn woodpecker. Scientific American 251(2): 114-121. Reprinted in: Behavior and Evolution of Birds. (D. W. Mock, ed.). W. H. Freeman, San Francisco.

  2. Koenig. W. D. 1988. La vie communicataire chez les oiseaux. La Recherche (Paris) 19: 300-308.

  3. Koenig, W. D. 1990. Oaks, acorns, and the acorn woodpecker. Fremontia 18: 77-79.

  4. Koenig, W. D. 1991. The acorn woodpecker. Pp. 86-87 in Oaks of California (Written by B. M. Pavlik, P. C. Muick, S. Johnson, and M. Popper). Cachuma Press, Los Olivos, CA.

  5. Koenig, W. D. 1992. [Defender of granaries: the acorn woodpecker]. Animals on Earth (Asahi Encyclopedia, weekly) 29: 137-139 [In Japanese].

  6. Koenig, W. D. 1993. Acorn woodpecker. Pp. 243-244 in The Marin County Breeding Bird Atlas: a Distributional and Natural History of Coastal California Birds, by W. D. Shuford. Bushtit Books, Bolinas, CA.

  7. Koenig, W. and J. Knops. 1995. Acorn production in California. Oaks ‘n’ folks 10(1): 1-4.

  8. Koenig, W. D. and J. Knops. 1995. Why do oaks produce boom-and-bust seed crops? California Agriculture 49(5): 7-12.

  9. Koenig, W. D. and R. L. Mumme. 1997. The great egg-demolition derby. Natural History 106(5): 32-37.

10. Koenig, W. and J. Knops. 1997. Geographical ecology of acorn production by California oaks. Oaks ‘n’ folks 12(2): 6-7.

11. Koenig, W. D. 1998. ‘Cooperative breeding, evolution of’ (pp. 159-160) and ‘ecological constraint hypothesis’ (p. 211) in The Encyclopedia of Ecology and Environmental Management. (P. Calow, editor-in-chief). Blackwell Publications, Oxford.

12. Kraaijeveld-Smit, F., and W. Koenig. 1998. Groepsgedrag van de Eikelspecht. Natura 1998(5): 140-141 [In Dutch].

13. Koenig, W. D. 2000. Oaks, acorns, and acorn woodpeckers. Oaks ‘n’ folks 15(1):5, 7.

14. Koenig, W. D. and J. Haydock. 2002. Group sex in the acorn woodpecker: who comes out on top, and why? The Biologist 49: 150-154. pdf

15. Dickinson, J. L. and W. D. Koenig. 2002. Social behaviour. Encyclopaedia Brittanica

16. Koenig, W. D. and J. M. H. Knops. 2005. The mystery of masting in trees. American Scientist 93: 340-347. pdf

17. Koenig, W. D. 2006. A look at Sudden Oak Death. BirdScope 20(2): 15.pdf

18. Dickinson, J. L. and W. D. Koenig. 2015. Social behavior, animal. Encyclopaedia Brittanica online; htty://www.britannica.com/EBchecked/topic/550897/animal-social-behaviour

19. Tietje, W. and W. Koenig. ND. Oaks, acorns, and acorn woodpecker. p. 8 in Wildlife among the oaks, written by S. G. Johnson with technical editing by W. D. Tietje. A publication of the Integrated Hardwoods Range Management Program

20. Koenig, W. 2011. Acorn woodpecker. p. 17 in The Monterey Pine Forest, by the monterey Pine Forest Watch. Pine Nut Press, Carmel-by-the-Sea, California.

21. Pearse, I. S. and W. D. Koenig. 2013. Acorns, acorns everywhere, but not in every year. International Oaks 24: 116-121.

22. Koenig, W. D. and J. L. Dickinson. 2014. Let's get together: the evolution of social behavior. Pp. 257-293 in Animal Behavior, vol. 2: Function and Evolution. K. Yasukawa, ed. Praeger, Santa Barbara, CA.

23. Koenig, W. D. 2016. Population biology. Pp. 495-534 in Handbook of Bird Biology, 3rd edition. I. Lovette and J. W. Fitzpatrick, eds. Cornell Lab of Ornithology, Ithaca, NY and Wiley, Hoboken, NJ.

24. Komdeur, J, and W. D. Koenig. 2016. Helpers and reproductive behavior in birds and mammals. In Elsevier Reference Module in Life Science, Encyclopedia of Animal Behavior. M. D. Breed and J. Moore, eds.

25. Koenig, W. D., M. B. Pesendorfer, and J. M. H Knops. 2013. Evolutionary drivers, proximate mechanisms, and spatial synchrony of acorn production in oaks. International Oaks 28: 29-40. pdf

26. Koenig, W. D. 2019. Acorn production and California oaks in a changing world. International Oaks 30: 45-52. pdf

27. Koenig, W. D. 2022. The acorn count: unraveling the mystery of oak tree mast years and their copious acorn crops is a big numbers game (with some weather thrown in). Bay Nature 22(3): 32-36. pdf

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Book reviews

 

 1. Koenig, W. D. 1983. Woodpeckers of the World, by L. L. Short. Wilson Bulletin 95: 698-700.

 2. Koenig, W. D. 1985. Perspectives in Ornithology, edited by A. H. Brush and G. A. Clark, Jr. American Scientist 73: 81-82.

 3. Koenig, W. D. 1985. The Florida Scrub Jay: Demography of a Cooperative-breeding Bird, by G. E. Woolfenden and J. W. Fitzpatrick. Animal Behaviour 33: 1402-1404.

 4. Koenig, W. D. 1986. The Life of the Woodpecker, by A. F. Skutch. Auk 103: 652-653.

 5. Koenig, W. D. 1987. Ecological Aspects of Social Evolution, edited by D. I. Rubenstein and R. W. Wrangham. Animal Behaviour 35: 1908-1909.

 6. Koenig, W. D. 1990. Reproductive Success. Studies of Individual Variation in Contrasting Breeding Systems, edited by T. H. Clutton-Brock. Auk 107: 223-224.

 7. Koenig, W. D. 1991. Food Hoarding in Animals, by S. B. Vander Wall. Journal of Evolutionary Biology 4: 690.

 8. Koenig, W. D. 1993. The Ostrich Communal Breeding System, by B. C. R. Bertram. Condor 95: 762-763.

 9. Koenig, W. D. 1994. The Natural History of Inbreeding and Outbreeding, edited by N. W. Thornhill. Journal of Wildlife Management 58: 797-798.

10. Koenig, W. D. 1996. Woodpeckers: a Guide to Woodpeckers of the World, by H. Winkler, D. A. Christie, and D. Nurney. Auk 113: 726-727.

11. Wolff, J. O. and W. D. Koenig. 1997. Cooperative Breeding in Mammals, edited by N. G. Solomon and J. A. French. Trends in Ecology & Evolution 12: 412-413.

12. Koenig, W. D. 1998. Made for Each Other, a Symbiosis of Birds and Pines, by R. N. Lanner. Quarterly Review of Biology 73: 100.

13. Koenig, W. D. 1999. Fairy-wrens and Grasswrens, by I. Rowley and E. Russell. Quarterly Review of Biology 74: 88-89.

14. Koenig, W. D. 2000. Helpers at Birds’ Nests: a Worldwide Survey of Cooperative Breeding and Related Behavior, by A. F. Skutch. Auk 117: 273-274.

15. Koenig, W. D. 2000. Hope is the Thing with Feathers, by C. Cokinos. Condor 102: 963-964.

16. Koenig, W. D. 2002. Dispersal, edited by J. E. Danchin, A. A. Dhondt, and J. D. Nichols. Ecology 83: 879-880.

17. Koenig, W. D. 2002. The Red-cockaded Woodpecker: Surviving in a Fire-maintained Ecosystem, by R. N. Conner, D. C. Rudolph, and J. R. Walters. Animal Behaviour 63: 831-832.

18. Koenig, W. D. 2010. Wytham Woods: Oxford’s Ecological Laboratory,, edited by P. S. Savill, C. M. Perrins, K. J. Kirby, and N. Fisher. Wilson Journal of Ornithology 22: 813-814.

19. Koenig, W. D. and E. L. Walters. 2014. Woodpeckers of the World: A Photographic Guide, by G. Gorman. Journal of Field Ornithology 85: 435-436.

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Obituaries

 

1. Koenig, W. D. and P. W. Sherman. 2004. In Memoriam: Frank Alois Pitelka, 1916-2003. International Society for Behavioral Ecology Newsletter 16: 4-5.

2. Koenig, W. D. and P. W. Sherman. 2004. In Memoriam: Frank Alois Pitelka, 1916-2003. Auk 121: 963-965.

3. Koenig, W. D. 2007. In Memoriam: Howard L. Cogswell, 1915-2006. Auk 124: 1096.

4. Koenig, W. D. and P. P. Marra. 2014. In Memoriam: Russell S. Greenberg, 1953-2013. Auk 131: 255-256.

4. Davis, W. E., Jr. and W. D. Koenig. 2019. In Memoriam: Thomas W. Custer, 1945-2019. Auk 136(4): ukz004.

 

Films and CD-ROMs

 

1. 1995. Featured in a 30-minute film on the ecology and social behavior of acorn woodpeckers produced by NHK, Japan, aired Oct. 1995.

2. 1997. Featured in Cal Alive! Exploring Biodiversity, an interactive educational CD-ROM about California biodiversity produced by the California Institute for Biodiversity.

3. 1998. For a few acorns more. National Geographic Explorer Series; produced and filmed by Oxford Scientific Films. 27 min film devoted to Koenig’s study of acorn woodpeckers. Originally aired 29 March 1998.

4. 1999. The Life of Birds . BBC; short section on acorn storage by acorn woodpeckers.

5. 2000. Birdwatch. PBS series; 9 min section on acorn woodpeckers and Koenig’s study.

6. 2003. Natural Reserve System: Hastings Reservation. A UCTV production airing nationally over cable TV featuring Hastings Reservation and including an extensive segment focusing on Koenig’s study of acorn woodpeckers. First aired 3 Dec. 2003. Available at http://www.uctv.tv/teachers/standards.asp?showID=8312

 

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Contact information:

Walter D. Koenig

38601 E. Carmel Valley Rd. Carmel Valley, CA 93924
Ph: 607 - 221-7591

wdk4@cornell.edu

 

Contact information:

Walter D. Koenig

Bird Population Studies
Cornell Lab of Ornithology
159 Sapsucker Woods Rd,
Ithaca, NY 14850
Office: 607 - 254-2151
wdk4@cornell.edu

My office in Mudd Hall: W307